Brassicaceae

I. tinctoria L., Sp. Pl., 670 (1753). Syn: I. canescens DC. in Lam. & DC. Fl. F?., 598 (1815)!
Biennial or shortly perennial. Stem 40-90 cm, glabrous or hirsute, subterete, laxly paniculate with branches coming off at more than 40° . Leaves hirsute. pubescent or glabrous, thin; basal leaves entire or weakly dentate; median stem leaves oblong or oblong-lanceolate with vertical triangular-lanceolate auricles. Flowers 3-4 mm, sepals glabrous. Fruits 10-20x2-5 mm. tapering from near apex to base, or oblong, glabrous or puberulent, wing thick, as wide or narrower than the median loculus (rarely wider), midrib slender. - A very polymorphic species of uncertain limits; the following subspecies may be provisionally recognised in Turkey:
1. Fruit 15-20 x 4-5 mm, gradually tapering from near apex to base, apex rounded or emarginate, glabrous, or with a puberulent loculus subsp. tinctoria
1. Fruit 10-15 x 2-4 mm
2. Fruit 12-15 x 2.5-4 mm, tapering from near apex to base, apex rounded or emarginate, usually puberulent-tomentose throughout subsp. tomentella
2. Fruit c. 10 x 2.5-3 mm, narrowly oblong, rounded at apex and base, glabrous or puberulent subsp. corymbosa
subsp. tomentella (Boiss.) Davis in Notes R.B.G. Edinb., 26: 22 (1964). Figure 16, p. 293. Syn: I. tinctoria var. stenocarpa Boiss., Fl. Or., 1: 381 (1867)! I. tomentella Boiss., Diagn., ser. 2(5): 46 (1856)! Fl. 5-8. Rocky slopes, scrub, waste places, 160-2400 m.
Type: [Turkey Bl Manisa] in regione media montis Sipyli supra Magnesiam Lydiae, Balansa 85 (K! W!).
Scattered in S. & E. Anatolia. B6 Seyhan: Amanus, c. Düldül, 1700-2300 m, Haradj. 2427! B8 Bingöl: Bingöl, 1800 m, Kotschy 1859: 754! B9 Bitlis: crater of Nemrut Da., 2400 m, McNeill 553! C4 Antalya: Alanya. 160 m, Hub.-Mor. 10448! C5 Niğde: Maaden, 1500 m, Siehe 1898: 167 (p.p.)! C9 Hakkari: Cilo Da. below Cilo Y., 2700 m, D. 24242!
Greece. Probably much commoner than the Turkish records suggest. Flower- , ing specimens from N. Anatolia may belong here. Alpine material from E. Anatolia is perennant and can be difficult to separate from I. takhtajanii (q.v.). Some Transcaucasian specimens referred by Avetisian (1963) to I. canescens DC. may represent I. tinctoria subsp. toinentella.